• Trinkle Professor of Biology

    Gilmer Hall, 107
    (434) 223-6326



Fulbright Scholar, Republic of Maldives, Spring 2006

Postdoctoral Research Fellow with Department of Wildlife Management, North Slope Borough, Barrow, Alaska

Ph.D. in organismic and evolutionary biology, Harvard University, 1992

A.M. in organismic and evolutionary biology, Harvard University, 1987

B.S. in zoology, Duke University, 1985, summa cum laude

Teaching Interests

Vertebrate functional morphology
Comparative anatomy and physiology
Evolutionary biology and paleontology
Marine biology and oceanography
Biomechanics and osteology
Mammalogy, ichthyology, and general vertebrate zoology
History and philosophy of science
Bioethics, human genomics, and human reproductive issues

Courses Taught

  • Vertebrate Anatomy
  • Vertebrate Physiology 
  • Evolutionary Theory/Population Genetics
  • Human Evolution/Physical Anthropology
  • Marine Biology
  • Evolutionary Ecology/Tropical Ecology (in Ecuador/Galapagos)
  • Organismal Biology
  • Principles of Biology
  • Vertebrate Paleontology
  • Physics of the Human Body (Biomechanics/Biomaterials)
  • Marine Mammals
  • Human Genome Project
  • Western Culture

Committee Assignments & Service to the College

Chair of Biology Department
Director of Honors Program and Chair of Honors Council
Health Science Advisory Committee and faculty advisor to PreHealth Society
Organizer of "Know Thyself: Human Genome Project" Symposium
Chair, Institutional Animal Care and Use Committee
President, Campus chapter of Phi Beta Kappa
President, Longwood/Hampden-Sydney chapter of Sigma Xi
Faculty rep, Duke Marine Laboratory/Marine Sciences Educational Consortium
Service on numerous other College committees (Faculty rep to Board of Trustees, President's Council, etc.)

Chair, Promotion & Tenure Committee
Chair, Academic Affairs Committee
Faculty Affairs Committee
Chair, Men's Studies Committee
Faculty oversight committee of Center for Entrepreneurship and Political Economy, and Wilson Center for Leadership in the Public Interest
Presidential Search Committee
Science Facility Steering Committee

Most Recent Peer-Reviewed Publications

[Werth's biomimetics research with a team of Dutch engineers using 3D printing to study filters has been profiled on wildinblue.com at www.wildinblue.com/featured.html]

Werth, A.J., S.M. Blakeney, and A.I. Cothren. 2019. Oil adsorption does not structurally or functionally alter whale baleen. Royal Society Open Science. http://dx.doi.org/10.1098/rsos.182194

Werth, A.J., M.A. Kosma, E.M. Chenoweth, and J.M. Straley. 2019. New views of humpback whale flow dynamics and morphology during prey engulfment. Marine Mammal Science. DOI: 10.1111/mms.12614

Werth, A.J., M.A. Lillie, M. Piscitelli, A.W. Vogl, and R.E. Shadwick. 2018. Slick, stretchy fascia underlies sliding tongue of rorquals. Anatomical Record 302:735-744 https://doi.org/10.1002/ar.24035

Werth, A.J., D. Rita Espada, M.V. Rosario, M.J. Moore, and T.L. Sformo. 2018. How do baleen whales stow their filter: a comparative biological analysis. Journal of Experimental Biology.  jeb.189233 doi: 10.1242/jeb.189233

Werth, A.J., J. Potvin, R.E. Shadwick, M.M. Jensen, D.E. Cade, and J.A. Goldbogen. 2018. Filtration area scaling and evolution in mysticetes: trophic niche partitioning and the curious cases of the sei and pygmy right whales. Biological Journal of the Linnean Society 125(2):264-279 doi/10.1093/biolinnean/bly121/5085357.

Werth, A.J., P. van de Graaf, and R. DesJardins. Preparation of full baleen racks for long-term exhibition and research. Provisionally accepted by Aquatic Mammals

Werth, A.J., C. Loch, and R.E. Fordyce. Evolution and devolution of odontocete enamel. Provisionally accepted by Journal of Mammalian Evolution.

Werth, A.J., and D. Allchin. How we think about human nature I: cognitive errors and concrete remedies. Provisionally accepted by Evolutionary Studies in Imaginative Culture

Allchin, D., and A.J. Werth. How we think about human nature II: the naturalizing error. Provisionally accepted by Philosophy of Science.

Werth, A.J. 2018. Variable porosity of throughput and tangential filtration in biological and 3D printed systems. In Advances in Engineering Research, Volume 29: Porosity—Properties and Measurement, ed. V. Petrova, pp. 1-58. Hauppauge, NY, Nova Science Publishers.

Werth, A.J. 2017. Are there limits to evolutionary explanations? Journal of Natural Sciences. 4(2):1-15.

Werth, A.J. 2017. Baleen. In: Encyclopedia of Marine Mammals 3e, ed. B. Wursig, J.G.M. Thewissen, K. Kovacs, pp 60-61. San Diego: Academic Press/Elsevier.

Werth, A.J., and H. Ito. 2017. Sling, scoop, squirter: anatomical features facilitating prey transport, concentration, and swallowing in rorqual whales (Mammalia: Mysticeti). Anatomical Record, doi: 10.1002/ar.23606.

Potvin, J., and A.J. Werth. 2017. Oral cavity hydrodynamics and drag production in balaenid whale suspension feeding. PLoS One (Public Library of Science), doi: 10.1371/journal.pone.0175220.

Goldbogen, J.A., D. Cade, J.A. Calambokidis, A.S. Friedlaender, J. Potvin, P.S. Segre, and A.J. Werth. 2017. How baleen whales feed: the biomechanics of engulfment and filtration. Annual Review of Marine Science, Vol. 9:367-386, doi 10.1146/annurev-marine-122414-033905.

Werth, A.J. 2017 [forthcoming, in press]. Baleen. In: Encyclopedia of Marine Mammals 3e, ed. J.G.M. Thewissen, B. Wursig, K. Kovacs. San Diego: Academic Press/Elsevier.

Werth, A.J., M.A. Lillie, M. Piscitelli, A.W. Vogl, and R.E. Shadwick. 2017 [in review]. Slick, stretchy fascia underlies sliding tongue or rorquals. Anatomical Record.

Werth, A.J., R. Harriss, M.V. Rosario, J.C. George, and T.L. Sformo. 2016. Hydration affects the physical and mechanical properties of baleen tissue. Royal Society Open Science 3: 160591, http://dx.doi.org/10.1098/rsos.160591 http://rsos.royalsocietypublishing.org/content/3/10/160591.abstract?ct=

Werth, A.J., and C. Loch. 2016 [in prep; to be submitted soon]. Phylogenetic and functional influences in the evolution and devolution of enamel in Cetacea. To be submitted to Mammalian Evolution.

Werth, A.J., M. Jensen, and J.A. Goldbogen. 2016 [in prep., to be submitted soon]. Filtration area, drag, and  the evolution of continuous vs. intermittent filtering strategies in Mysticeti. To be submitted to Journal of Mammalogy.

Werth, A.J., and J. Potvin. 2016. Baleen hydrodynamics and morphology of crossflow filtration in balaenid whale suspension feeding. PLOS One, doi:10.1371/journal.pone.01501016. http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0150106

Werth, A.J., J. Straley, and R.E. Shadwick. 2016. Baleen wear reveals intraoral water flow patterns of mysticete filter feeding. Journal of Morphology, doi:10.1002/jmor.20510. http://onlinelibrary.wiley.com/doi/10.1002/jmor.20510/abstract

Allchin, D.A., and A.J. Werth. 2016. The naturalizing error: implications for appeals to human nature and self-justifying nature. Accepted by Journal for General Philosophy of Science.

Werth, A.J. 2015. Inertia vs. freedom in faculty life: Surviving a clash between the most powerful force in the universe and the most perilous perk in academe. The Chronicle of Higher Education August 17, 2015. http://chromicle.com/article/Inertia-vs-Freedom-in-Faculty/232435/?cid=at&utm_source=at&utm_medium=en http://chronicle.com/article/Inertia-vs-Freedom-in-Faculty/232435/

Werth, A.J., and W.A. Shear. 2014. The evolutionary truth about living fossils. American Scientist 102(6):434-443. doi: 10.1511/2014.111.434.

Werth, A.J. 2014. What makes biology tick? Review of In Search of Mechanisms: Discoveries Across the Life Sciences book by Carl F. Craver and Lindley Darden. BioScience 64(9):845-847.

Werth, A.J. 2014. Vestiges of the natural history of development: historical holdovers reveal the dynamic interaction between ontogeny and phylogeny. Evolution: Education and Outreach 7:12:1-11. doi: 10.1186/s12052-014-0012-5. http://link.springer.com/article/10.1186%2Fs12052-014-0012-5

Werth, A.J. 2014. Why single-sex education is a viable educational option. In What Works: Raising Boys, Engaging Guys, Educating Men. Hampden-Sydney College.

Werth, A.J. 2013. Flow-dependent porosity of baleen. Journal of Experimental Biology 216:1152-1159. doi:10.1242/jeb.078931.Ford, T.J., A.J. Werth, and J.C. George. 2013. An intraoral thermoregulatory organ in the bowhead whale (Balaena mysticetus), the corpus caervosum maxillaris. Anatomical Record: Advances in Integrative Anatomy and Evolutionary Biology 296:701-708. doi:10.1002/ar.22681.

Werth, A.J. 2013. An evolutionary focus improves students' understanding of all biology.  Reports of the National Center for Science Education 33(1):1-18, http://reports.ncse.com/index.php/rncse/article/view/31/251.

Werth, A.J. 2012. Hydrodynamic and sensory factors governing response of copepods to simulated predation by baleen whales. International Journal of Ecology. http://www.hindawi.com/journals/ijeco/2012/208913/

Werth, A.J., and T.J. Ford. 2012. Abdominal fat pads act as control surfaces in lieu of dorsal fins in the beluga (Delphinapterus). Marine Mammal Science 28(4):E516-527. doi:10.1111.j.1748-7692.2012.00567.x.

Werth, A.J. 2012. Avoiding the pitfall of progress and asociated perils of evolutionary education. Evolution: Education and Outreach 5(2)249-265. doi:10.1007/s120052-012-0417-7.

Werth, A.J. 2012. From fins to legs and back again. Review of Return to the Sea: The Life and Evolutionary Times of Marine Mammals book by Annalisa Berta. BioScience 62(11):996-997.

Allchin, D., and A.J. Werth [forthcoming] The naturalizing error: Implications for appeals to human nature and self-justifying nature. In peer review.

Werth. A.J. [in prep] Cross-flow filtration in baleen. For submission to Journal of Morphology.Werth, A.J. [forthcoming] Huxley's garden, Wallace's ghost, and Harlow's brain: Are there limits to evolutionary explanation? In preparation; for submission to Zygon.Beatty, B., and A.J. Werth [forthcoming] Aprismatic enamel microstructure and tooth sharpness in odontocetes: Economy or adaptation? In preparation; for submission to Integrative Biology.

Werth, A.J. 2009. Clearing the highest hurdle:Human-based case studies broaden students' understanding of core evolutionary concepts. The Journal of Effective Teaching, 9(2): 37-52.

Werth, A.J. 2008. The human genome project: implications for the study of human evolution. In The Human Genome Project: Ethical, Legal, and Social Implications. University Press of New England.

Werth, A.J. 2007. Adaptations of the cetacean hyolingual apparatus for aquatic feeding and thermoregulation. The Anatomical Record, 290(6): 546-568.Werth, A.J. 2006. Odontocete suction feeding: Experimental analysis of water flow and head shape. Journal of Morphology, 267(12): 1415-1428.

Werth, A.J. 2006. Mandibular and dental variation and the evolution of suction feeding in Odontoceti. Journal of Mammalogy, 87(3): 579-588.

Werth, A.J. 2005. Evolution and development are not the same. The American Biology Teacher, 67(4): 201-202.Werth, A.J. 2005. On the benefits of teaching honors. Honors In Practice [The National Honors Report], 1(1):43-48.

Werth, A.J. 2004. Functional morphology of the sperm whale tongue, with reference to suction feeding. Aquatic Mammals 30(3), 405-418.

Werth, A.J. 2004. Models of hydrodynamic flow in the bowhead whale filter feeding apparatus. Journal of Experimental Biology 207(20), 3569-3580. http://jeb.biologists.org/content/207/20/3569.full

Werth, A.J. 2003. Unity in diversity: The virtues of a metadisciplinary perspective in liberal arts education. Journal of the National Collegiate Honors Council, Fall/Winter 2003: 35-52.

Werth, A.J. 2001. How do mysticetes remove prey trapped in baleen? Bulletin of the Museum of Comparative Zoology, 156(1): 189-203.

Werth, A.J. 2001. Simple lessons in biomechanics using everyday objects. The American Biology Teacher, 63(4): 267-270.

Werth, A.J. 2000. Marine Mammals. In Feeding: Form, Function and Evolution in Tetrapod Vertebrates, ed. K. Schwenk, pp. 475-514. New York, Academic Press.

Werth, A.J. 2000. A kinematic study of suction feeding and associated behaviors inthe long-finned pilot whale, Globicephala melas. Marine Mammal Science, 16(2): 299-314.In addition to peer-reviewed publications, Werth's research has been profiled by journalists and science writers in print and online as well as via other media (radio, television, blogs, etc.), including but not limited to the following: Science, National Geographic, New Scientist, Science News, Live Science, Discover Magazine, Discovery Canada, Scientific American, NBC, BBC, LeMonde, Sciences Avenir: Le Nouvel Observateur, French Thermography Association, CBC (Quirks and Quarks national radio program (www.cbc.ca/quirks/), Swedish National Radio.Here is a sampling of such references:Davies, E. 2013. Whales filter feed with a tangled hair-like net. BBC Nature Newshttp://www.bbc.co.uk/nature/21755885

Lewis, T. 2013. Whales trap dinner with mouthful of swirling bristles. Live Science http://www.livescience.com/27878-how-whale-baleen-traps-food.htmlSmith, B. 2013. Whales' baleen entangle prey with hairy bristles. Red Orbithttp://www.redorbit.com/news/science/1112803585/baleen-whale-teeth-entangle-prey-031413/

Knight, K. 2013. Streaming baleen tangles to trap food. "Inside JEB" commentary article written about Werth's featured paper in Journal of Experimental Biologyhttp://jeb.biologists.org/content/216/7/i.2

Zimmer, C. 2013. The brain-chilling, shrimp-caressing, lamppost-sized, NSFW organ hiding in a whale's mouth. National Geographic: The Loom blog entry (posted March 5, 2013) about Werth's research article published in The Anatomical Record. http://phenomena.nationalgeographic.com/2013/03/04/the-brain-chilling-shrimp-caressing-lamppost-sized-nsfw-organ-hiding-in-a-whales-mouth/

Marshall, M. 2013. World's biggest mouth has an erectile secret. New Scientisthttp://www.newscientist.com/article/dn23252-zoologger-worlds-biggest-mouth-has-an-erectile-secret.html

McDonald, B. 2013. Penis-like organ helps bowheads keep cool. Quirks & Quarks CBC radio program broadcast, April 20, 2013. http://www.cbc.ca/quirks/episode/2013/04/20/april-20-2013/index.html#2

Pennisi, E. 1999. Coming to grips with sperm whale anatomy. Science 283(5401):475-477.Svitil, K. 1999. The table manners of whales. Discover 20(5): 20.

Honors & Awards

Phi Beta Kappa, Omicron Delta Kappa

Fulbright Scholar (teaching/research), Republic of Maldives: "Marine Environmental Science of Maldive Islands"

2014 NSF-funded Scientist in Residency Fellow, Sitka Sound Science Center, Sitka, Alaska

2013 Visiting Professor of Zoology, University of British Columbia

2012 awarded Charles Scott Venable endowed Professorship

2011 Harris Teaching Award from VFIC:http://www.hsc.edu/News/News-Archive/2011/Werth.htmlhttp://www.vfic.org/programs/harris_index.html

Invited to Smithsonian/NESCent workshops on Teaching Human Evolution

2010 Mettauer Research Award, H-SC
2008 Cabell Teaching Award, H-SC
2006 Fuqua Teaching Award, H-SC
2001 Crawley Teaching Award, H-SC
2002 Wye Faculty Seminar Fellow
2000 NIH Faculty Summer Fellow, Dartmouth College
1996 VFIC Mednick Fellow (research funding)
Visiting Scholar, University of Maine Darling Marine Center
Visiting Scholar, Providence College
Research funding from government of Alaska
Research funding from H-SC faculty fellowships

Professional Affiliations

Sigma Xi (Scientific Research Society)
National Association of Biolgoy Teachers, Virginia Association of Biology Teachers
National Collegiate Honors Council, Southern Regional Honors Council, Virginia Collegiate Honors Council
Society for Integrative and Comparative Biology
Society for Marine Mammalogy
International Society of Vertebrate Morphology
Society for the Study of Mammalian Evolution
Virginia Natural History Society

Reviewer for NSF and 20 journals

Other Interests

running & hiking
reading & drawing
SCUBA & snorkeling
listening to music
playing & watching sports
ardent Duke basketball and (no longer!) condemned Red Sox fan

wife Diane
son Colin
daughter Gwen

Research Interests and potential topics for students

Research interests: specifically, structure & function of marine mammals, especially feeding in toothed and baleen whales; biomechanics & materials science of biological tissues; 3D printing of biological structures or biological analogues; basic fluid dynamics and effects of flow on organisms; fossils & paleontology
Potential topics for students: marine biology & oceanography; anatomy & physiology; anything related to evolution (human and otherwise); science & religion; philosophy of science; bioethics; environmental issues